Wilaru is an extinct genus of presbyornithid bird from Australia during the Late Oligocene to Early Miocene, around 26-22 million years ago.[2] The type species is Wilaru tedfordi, and the second species is Wilaru prideauxi.[3]
Discovery and naming
![](https://upload.wikimedia.org/wikipedia/commons/thumb/7/71/Postcranial_elements_of_Presbyornithidae_and_southern_screamer.jpg/220px-Postcranial_elements_of_Presbyornithidae_and_southern_screamer.jpg)
The type species, Wilaru tedfordi, was described from fossil material collected from Lake Pinpa, Lake Palankarinna and Billeroo Creek, specfically in the Namba Formation of Lake Eyre Basin, north-eastern South Australia. The genus name Wilaru is the term for “stone curlew” in the Diyari language of the Lake Eyre region. The specific epithet of the type species W. tedfordi honours American palaeontologist Richard H. Tedford (1929–2011) of the American Museum of Natural History, who led the 1971 expedition to Lake Pinpa during which much of the descriptive material was collected.[4]
The second species, Wilaru prideauxi, was first named in 2016 after the Australian vertebrate paleontologist Gavin Prideaux, who worked on Lake Ngapakaldi of the Wipajiri Formation in South Australia where the specimen of W. prideauxi was discovered and on various Australian mammals which lived during the Oligocene to Miocene epochs.[3] W. prideauxi was properly named in 2020 after the authors republished the study as a correction with Zoobank Registration.[1][2] The holotype of W. tedfordi is a left humerus (SAMA P48925), while the holotype of W. prideauxi is a right tarsometatarsus (SAMA P53136).[3]
Description and ecology
Compared to other presbyornithids, Willaru appeared to have been specialised to a more terrestrial lifestyle, based on its tarsometatarsal morphology. In particular, the latter W. prideauxi appears to have been more specialised towards terrestriality than the earlier W. tedfordi, being larger and more robust, indicating a clear speciation towards this lifestyle and therefore a direct species sequence.[3]
Like many modern waterfowl, the Willaru species had spurs and knobs on their carpals. Like the closely related modern-day screamers, these were almost certainly used to fight, indicating perhaps territorial habits, as opposed to the more gregarious nature of earlier presbyornithids.[3]
Willaru co-existed with several anatid and anseranatid species, indicating that there was little ecological competition. It is possible that a speciation towards terrestriality might have spared it from competition with more derived waterfowl, allowing it to live longer than other presbyornithids.[3]
Classification
It was originally classified as a stone-curlew,[4] but subsequently it was argued to be the youngest member of the extinct family Presbyornithidae instead.[3] The discovery of a similar Eocene presbyornithid Murgonornis archeri also supported this taxonomic assignment.[2] A 2024 phylogenetic analysis recovered Wilaru within various positions of Galloanserae, making its identity as a presbyornithid questionable.[5] However, a 2025 study which reported a nearly complete skull of Vegavis revised the phylogenetic analyses of Galloanserae and recovered Wilaru as a member of the Presbyornithidae within Anseriformes.[6]
References
- ^ a b De Pietri, V.L.; Scofield, R.P.; Zelenkov, N.; Boles, W.E.; Worthy, T.H. (2020). "Correction to 'The unexpected survival of an ancient lineage of anseriform birds into the Neogene of Australia: the youngest record of Presbyornithidae'". Royal Society Open Science. 7 (11). 201430. doi:10.1098/rsos.201430. ISSN 2054-5703. PMC 7735352. PMID 33391810.
- ^ a b c Worthy, Trevor H.; De Pietri, Vanesa L.; Scofield, R. Paul; Hand, Suzanne J. (2023-03-20). "A new Eocene species of presbyornithid (Aves, Anseriformes) from Murgon, Australia". Alcheringa: An Australasian Journal of Palaeontology. 47 (4): 416–430. Bibcode:2023Alch...47..416W. doi:10.1080/03115518.2023.2184491. hdl:1959.4/unsworks_83108. ISSN 0311-5518. S2CID 257679005.
- ^ a b c d e f g De Pietri, V.L.; Scofield, R.P.; Zelenkov, N.; Boles, W.E.; Worthy, T.H. (2016). "The unexpected survival of an ancient lineage of anseriform birds into the Neogene of Australia: the youngest record of Presbyornithidae". Royal Society Open Science. 3 (2). 150635. Bibcode:2016RSOS....350635D. doi:10.1098/rsos.150635. ISSN 2054-5703. PMC 4785986. PMID 26998335.
- ^ a b Walter E. Boles; Melanie A. Finch; Rene H. Hofheins; Patricia Vickers-Rich; Mary Walters & Thomas H. Rich (2013). "A fossil stone-curlew (Aves: Burhinidae) from the Late Oligocene/Early Miocene of South Australia". In Ursula B. Göhlich & Andreas Kroh (eds.). Paleornithological Research 2013. Proceedings of the 8th International Meeting of the Society of Avian Paleontology and Evolution, Vienna, 2012 (PDF). Naturhistorisches Museum Wien. pp. 43–61. ISBN 978-3-902421-82-1.
- ^ Musser, G.; Clarke, J. A. (2024). "A new Paleogene fossil and a new dataset for waterfowl (Aves: Anseriformes) clarify phylogeny, ecological evolution, and avian evolution at the K-Pg Boundary". PLOS ONE. 19 (7). e0278737. doi:10.1371/journal.pone.0278737. PMC 11288464. PMID 39078833.
- ^ Torres, Christopher R.; Clarke, Julia A.; Groenke, Joseph R.; Lamanna, Matthew C.; MacPhee, Ross D. E.; Musser, Grace M.; Roberts, Eric M.; O’Connor, Patrick M. (2025). "Cretaceous Antarctic bird skull elucidates early avian ecological diversity". Nature. 638 (8049): 146–151. doi:10.1038/s41586-024-08390-0. ISSN 1476-4687.
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