The Arcillas de Morella Formation is a geological formation in Spain whose strata date back to the Barremian stage of the Early Cretaceous.[1] Dinosaur remains are among the fossils that have been recovered from the formation.[2]

Geology

The formation was formally named and defined by Canérot and colleagues in 1982.[3] The formation predominantly consists of red clays and continental sandstones, with some marine intercalations.[4]

Vertebrate paleofauna

Dinosaurs of the Arcillas de Morella Formation
Taxa Species Notes Images
Hypsilophodon[2]
Hypsilophodon
Iguanodon
Mantellisaurus
Protathlitis
Vallibonavenatrix
Iguanodon I. bernissartensis[5] An iguanodonian dinosaur
Mantellisaurus M. cf. atherfieldensis[5] An iguanodontian dinosaur
Morelladon Morelladon beltrani[6] An iguanodontian dinosaur
Baryonychinae[2] Indeterminate remains[2]
cf. Carcharodontosauridae Indeterminate remains[7] An anterior caudal centrum belonging to an immature individual
Garumbatitan G. morellensis A somphospondylan sauropod[8]
Polacanthus P. sp A polacanthine nodosaurid ankylosaur
Protathlitis P. cinctorrensis[9] A spinosaurid theropod
Titanosauriformes Indeterminate At least two distinct taxa, including a somphospondylan[10]
Vallibonavenatrix V. cani[4] A spinosaurid theropod

Turtles

Turtles of the Arcillas de Morella Formation
Taxa Species Notes Images
Brodiechelys B. royoi[11] A xinjiangchelyid turtle
Eodortoka E. morellana[12] A dortokid turtle

Plesiosaurs

Plesiosaurs of the Arcillas de Morella Formation
Taxa Presence Notes Images
Leptocleididae[13]

Correlation

Early Cretaceous stratigraphy of Iberia
Ma Age Paleomap \ Basins Cantabrian Olanyà Cameros Maestrazgo Oliete Galve Morella South Iberian Pre-betic Lusitanian
100 Cenomanian
La Cabana Sopeira Utrillas Mosquerela Caranguejeira
Altamira Utrillas
Eguino
125 Albian Ullaga - Balmaseda Lluçà Traiguera
Monte Grande Escucha Escucha Jijona
Itxina - Miono
Aptian Valmaseda - Tellamendi Ol Gp. - Castrillo Benassal Benassal Olhos
Font En Gp. - Leza Morella/Oliete Oliete Villaroya Morella Capas
Rojas
Almargem
Patrocinio - Ernaga Senyús En Gp. - Jubela Forcall Villaroya Upper
Bedoulian
Figueira
Barremian Vega de Pas Cabó Abejar Xert Alacón Xert Huérguina Assises
Prada Artoles Collado Moutonianum Papo Seco
Rúbies Tera Gp. - Golmayo Alacón/Blesa Blesa Camarillas Mirambel
150 Hauterivian Ur Gp. - Pinilla Llacova Castellar Tera Gp. - Pinilla Villares Porto da
Calada
hiatus
Huerva Gaita
Valanginian Villaro Ur Gp. - Larriba Ped Gp. - Hortigüela
Ped Gp. - Hortigüela Ped Gp. - Piedrahita
Peñacoba Galve Miravetes
Berriasian Cab Gp. - Arcera Valdeprado hiatus Alfambra
TdL Gp. - Rupelo Arzobispo hiatus Tollo
On Gp. - Huérteles
Sierra Matute
Tithonian Lastres Tera Gp. - Magaña Higuereles Tera Gp. - Magaña Lourinhã
Arzobispo
Ágreda
Legend Major fossiliferous, oofossiliferous, ichnofossiliferous, coproliferous, minor formation
Sources


See also

References

  1. ^ Villanueva-Amadoz, Uxue; Santisteban, Carlos; Santos-Cubedo, Andrés (2015-04-03). "Age determination of the Arcillas de Morella Formation (Maestrazgo Basin, Spain)". Historical Biology. 27 (3–4): 389–397. Bibcode:2015HBio...27..389V. doi:10.1080/08912963.2013.874422. ISSN 0891-2963. S2CID 128775513.
  2. ^ a b c d Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 562. ISBN 0-520-24209-2.
  3. ^ Villanueva-Amadoz, Uxue; Santisteban, Carlos; Santos-Cubedo, Andrés (2015-04-03). "Age determination of the Arcillas de Morella Formation (Maestrazgo Basin, Spain)". Historical Biology. 27 (3–4): 389–397. Bibcode:2015HBio...27..389V. doi:10.1080/08912963.2013.874422. ISSN 0891-2963. S2CID 128775513.
  4. ^ a b Elisabete Malafaia; José Miguel Gasulla; Fernando Escaso; Iván Narváez; José Luis Sanz; Francisco Ortega (2019). "A new spinosaurid theropod (Dinosauria: Megalosauroidea) from the late Barremian of Vallibona, Spain: Implications for spinosaurid diversity in the Early Cretaceous of the Iberian Peninsula". Cretaceous Research. in press: 104221. doi:10.1016/j.cretres.2019.104221.
  5. ^ a b Gasulla, José M.; Escaso, Fernando; Ortega, Francisco; Sanz, José L. (January 2014). "New hadrosauriform cranial remains from the Arcillas de Morella Formation (lower Aptian) of Morella, Spain". Cretaceous Research. 47: 19–24. Bibcode:2014CrRes..47...19G. doi:10.1016/j.cretres.2013.10.004.
  6. ^ Gasulla, José Miguel; Escaso, Fernando; Narváez, Iván; Ortega, Francisco; Sanz, José Luis (2015). "A New Sail-Backed Styracosternan (Dinosauria: Ornithopoda) from the Early Cretaceous of Morella, Spain". PLOS ONE. 10 (12): e0144167. Bibcode:2015PLoSO..1044167G. doi:10.1371/journal.pone.0144167. PMC 4691198. PMID 26673161.
  7. ^ Montealegre, Adrian; Castillo-Visa, Oscar; Sellés, Albert (2024-01-31). "New theropod remains from the late Barremian (Early Cretaceous) of Eastern Iberian Peninsula". Historical Biology: 1–11. doi:10.1080/08912963.2024.2308220. ISSN 0891-2963.
  8. ^ Mocho, Pedro; Escaso, Fernando; Gasulla, José M; Galobart, Àngel; Poza, Begoña; Santos-Cubedo, Andrés; Sanz, José L; Ortega, Francisco (2023-09-28). "New sauropod dinosaur from the Lower Cretaceous of Morella (Spain) provides new insights on the evolutionary history of Iberian somphospondylan titanosauriforms". Zoological Journal of the Linnean Society. 201: 214–268. doi:10.1093/zoolinnean/zlad124. ISSN 0024-4082.
  9. ^ Santos-Cubedo, A.; de Santisteban, C.; Poza, B.; Meseguer, S. (2023). "A new spinosaurid dinosaur species from the Early Cretaceous of Cinctorres (Spain)". Scientific Reports. 13 (1). 6471. Bibcode:2023NatSR..13.6471S. doi:10.1038/s41598-023-33418-2. hdl:10234/203142. PMID 37202441.
  10. ^ Mocho, P.; Pérez-García, A.; Gasulla, J. M.; Ortega, F. (2017-06-01). "High sauropod diversity in the upper Barremian Arcillas de Morella Formation (Maestrat Basin, Spain) revealed by a systematic review of historical material". Journal of Iberian Geology. 43 (2): 111–128. Bibcode:2017JIbG...43..111M. doi:10.1007/s41513-017-0012-8. ISSN 1886-7995. S2CID 134844724.
  11. ^ Pérez-García, A.; Gasulla, J.M.; Ortega, F. (2014). "A new turtle species of Brodiechelys from the Early Cretaceous of Spain: Systematic and palaeobiogeographic implications" (PDF). Acta Palaeontologica Polonica. 59 (2): 333–342.
  12. ^ Pérez-García, A.; Gasulla, J.M.; Ortega, F. (2014-03-01). "Eodortoka morellana gen. et sp. nov., the first pan-pleurodiran turtle (Dortokidae) defined in the Lower Cretaceous of Europe". Cretaceous Research. 48: 130–138. Bibcode:2014CrRes..48..130P. doi:10.1016/j.cretres.2013.12.004. ISSN 0195-6671.
  13. ^ Quesada, Juan M.; Pérez-García, Adán; Gasulla, José Miguel; Ortega, Francisco (2019-02-01). "Plesiosauria remains from the Barremian of Morella (Castellón, Spain) and first identification of Leptocleididae in the Iberian record". Cretaceous Research. 94: 8–24. Bibcode:2019CrRes..94....8Q. doi:10.1016/j.cretres.2018.10.010. ISSN 0195-6671. S2CID 134139253.

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