This list of informally named Mesozoic reptiles is a listing of prehistoric reptiles from the Mesozoic era (excluding dinosaurs and pterosaurs) that have never been given formally published scientific names. This list only includes names that were not properly published ("unavailable names") and have not since been published under a valid name. The following types of names are present on this list:
- Nomen nudum, Latin for "naked name": A name that has appeared in print but has not yet been formally published by the standards of the International Commission on Zoological Nomenclature. Nomina nuda (the plural form) are invalid, and are therefore not italicized as a proper generic name would be.
- Nomen manuscriptum, Latin for "manuscript name": A name that appears in manuscript but was not formally published. A nomen manuscriptum is equivalent to a nomen nudum for everything except the method of publication, and description.
- Nomen ex dissertationae, Latin for "dissertation name": A name that appears in a dissertation but was not formally published.
- Nicknames or descriptive names given to specimens or taxa by researchers or the press.
A
Aust Colossus
The "Aust Colossus" is an informal name given to the giant ichthyosaur specimens (BRSMG Cb3869, BRSMG Cb3870, BRSMG Cb4063 and BRSUG 7007) discovered from the Late Triassic (Rhaetian) Westbury Formation at the Aust Cliff.[1][2] "Aust Colossus" has been tentatively estimated to be even larger than the 25 metres (82 ft) long ichthyosaur Ichthyotitan from the same formation, possibly over 30 metres (98 ft) long, though the authors acknowledge that this is a very speculative estimate.[3]
C
Cystosaurus
"Cystosaurus" is a genus of putative teleosaur from France informally named by Étienne Geoffroy Saint-Hilaire in 1831.[4] Because it was never given formal description, "Cystosaurus" remains a nomen nudum.[5] Cryptodraco was used as an unnecessary replacement name for Cryptosaurus by Richard Lydekker (1889) who believed the name Cryptosaurus was already previously in use for "Cystosaurus".[6]
H
Hadongsuchus
"Hadongsuchus" (crocodile of Hadong County) is an Early Cretaceous (Aptian-Albian) crocodyliform from the Hasandong Formation of South Korea. Known from a 52 mm (2.0 in) long complete skull (KIGAM VP 200401) discovered in 2002, it was informally named as "Hadongsuchus acerdentis" in a 2005 thesis. Certain features of the third premaxillary tooth help distinguish it from other related crocodylomorphs. Like other protosuchians, it is believed to have been a fully terrestrial cursorial animal with a semi-erect posture. Lee (2005) suggested that "Hadongsuchus" and three other crocodyliforms (Shantungosuchus, Sichuanosuchus, and Zosuchus) comprise a new family which represents a sister taxa to the Protosuchidae, with Zosuchus being the closest relative.[7]
Hainosaurus boubker
Hainosaurus "boubker" is an informal species of mosasaur from the Sidi Chennane phosphate quarry in Morocco.[8] The description of H. "boubker" was published by a known predatory journal, Scientific Research Publishing, which places the validity of the publication and this taxon in questionable status.[9][10] Some researchers provisionally considered H. "boubker" as a valid taxon.[11]
I
Induszalim
"Induszalim" is an informal genus of crocodyliform named by M.S. Malkani based on fragmentary remains from the Vitakri Formation of Pakistan. The proposed type species, "Induszalim bala", was named in a 2014 conference abstract, rendering this name a nomen nudum.[12]
K
Khuzdarcroco
"Khuzdarcroco" is an informal genus of purported mesoeucrocodylians proposed by M.S. Malkani in a 2015 conference abstract based on a rib fragment from the Goru Formation of Pakistan. The proposed type species is, "Khuzdarcroco zahri".[13]
M
Monster of Aramberri
The "Monster of Aramberri", also known as the "Aramberri pliosaur", is an informal name given to a large pliosaur specimen (UANL-FCT-R2) discovered from the Late Jurassic (Kimmeridgian) La Caja Formation, Mexico.[14] The body length of the "Aramberri pliosaur" is estimated around 10–11 metres (33–36 ft).[15]
N
Narynsuchus
"Narynsuchus" is an extinct genus of goniopholidid. Because it has never been formally described, it is currently considered a nomen nudum. The name was first used in 1990 for fossil material found from a formation called the Balabansai Svita, which forms a lens in site FTA-30 of the Sarykamyshsai 1 locality in the Fergana Valley of Kyrgyzstan.[16] The formation dates back to the Callovian stage of the Middle Jurassic.[17] The known material of the intended type species "Narynsuchus ferganensis" consists of large and characteristically striated teeth as well as a robust left tibia.[18] "Narynsuchus" is thought to have been larger than Sunosuchus, another crocodyliform from Sarykamyshsai 1, due to the greater size of the known elements belonging to the genus.[17] This material was first recognized as belonging to a new crocodyliform in 1989, when it was referred to as an indeterminant species of Peipehsuchus.[18]
P
Platychelone
"Platychelone" is an informal genus of sea turtle from the Maastrichtian type area of Netherlands. Named by Louis Dollo in 1909, the intended type species "Platychelone emarginata" is considered a nomen nudum as the taxon was based on an undescribed, large carapace with no illustrations.[19]
Procoelosaurus
"Procoelosaurus" is an informal genus of purported pterosauromorph from the Late Triassic Cooper Canyon Formation of Texas. Named in a 2002 thesis, the intended type species "Procoelosaurus brevicollis" was classified as an ornithodiran close to pterosaurs based on its hindlimb morphology.[20] Other potential specimens identified as cf. "Procoelosaurus" were noted in a 2008 thesis.[21]
Pteromimus
"Pteromimus" is an informal genus of purported pterosauromorph from the Late Triassic Cooper Canyon Formation of Texas. Named in a 2002 thesis, the intended type species "Pteromimus longicollis" was classified as an ornithodiran close to pterosaurs based on its hindlimb morphology.[20] A cervical vertebra identified as cf. "Pteromimus" was noted in a 2008 thesis.[21]
R
Raptocleidus
"Raptocleidus" is a plesiosaur from the Early Jurassic (Pliensbachian) deposits of Lyme Regis, southern United Kingdom. The genus contained two species, "Raptocleidus blakei" (LEICT G1.2002) and "Raptocleidus bondi" (NHMUK R16330), both of which were informally named in a 2012 thesis. It was likely a small plesiosaur, with an estimated body length of 3 metres (9.8 ft).[22]
Richmond pliosaur
The "Richmond pliosaur", also known as "Penny", is an informal name given to a complete polycotylid specimen (QM F18041) from the Early Cretaceous (Albian) Allaru Formation, Australia.[23][24] Discovered in October 1989, this complete articulated specimen is measured around 5 metres (16 ft) long.[25] Within the family Polycotylidae, it is classified as a member of the clade Occultonectia, which also includes Plesiopleurodon and Sulcusuchus.[26]
S
Satsuma-utsunomiya-ryu

"Satsuma-utsunomiya-ryu" is the nickname given to an elasmosaurid plesiosaur from Kyushu, Japan. It is the oldest known elasmosaurid from East Asia, dating back to approximately 100 million years ago. The known material includes the mandible, hyoid, fragments of the skull, 40 cervical vertebrae, and a limb bone, possibly belonging to a juvenile individual. Features of the tooth surface ornamentation cervical vertebrae, suggest Satsuma-unomiya-ryu may represent a new species. The name references the former province name of the discovery site and the discoverer, Satoshi Utsunomiya.[27][28] In 2021, Ustunomiya and Yasuhisa Nakajima reported the presence of a regurgitated "pellet" around its throat region.[29]
Stereosaurus
"Stereosaurus" is a plesiosaur from the Cambridge Greensand (Cenomanian). The genus contains three species, "Stereosaurus platyomus", "S. cratynotus" and "S. stenomus", all of which were informally coined in 1869 by British paleontologist Harry Seeley, who considered them to be plesiosaurian.[30][31]
Sulaimanisuchus
"Sulaimanisuchus" is an informal genus of purported mesoeucrocodylians from the Cretaceous Vitakri Formation of Pakistan, mentioned by M. Sadiq Malkani in 2010. The proposed holotype is a partial mandibular symphysis. The intended type species is "Sulaimanisuchus kinwai."[32]
T
Tylosaurus borealis
Tylosaurus "borealis" is an informal species of mosasaur, with the known specimen discovered from approximately 55 kilometers (34 mi) northeast of Grande Prairie. It was named in a 2020 thesis by Samuel Garvey who described a partial skull of Tylosaurus catalogued as TMP 2014.011.0001, which makes the specimen the northernmost known occurrence of this genus.[33]
Z
Zahrisaurus
"Zahrisaurus" is an informal genus of purported plesiosaurs from the ?Jurassic Sulaiman Group of Balochistan, described by M. Sadiq Malkani (2019) in Scientific Research Publishing, a known predatory publisher. The proposed holotype is allegedly a portion of the trunk with associated ribs. The intended type species is "Zahrisaurus kilmoolai."[34]
See also
References
- ^ Gayford, Joel H.; Engelman, Russell K.; Sternes, Phillip C.; Itano, Wayne M.; Bazzi, Mohamad; Collareta, Alberto; Salas-Gismondi, Rodolfo; Shimada, Kenshu (September 2024). "Cautionary tales on the use of proxies to estimate body size and form of extinct animals". Ecology and Evolution. 14 (9): e70218. doi:10.1002/ece3.70218. ISSN 2045-7758. PMC 11368419. PMID 39224151.
- ^ Lomax, Dean R.; De la Salle, Paul; Massare, Judy A.; Gallois, Ramues (9 April 2018). Wong, William Oki (ed.). "A giant Late Triassic ichthyosaur from the UK and a reinterpretation of the Aust Cliff 'dinosaurian' bones". PLOS ONE. 13 (4): e0194742. Bibcode:2018PLoSO..1394742L. doi:10.1371/journal.pone.0194742. ISSN 1932-6203. PMC 5890986. PMID 29630618.
- ^ Lomax, D. R.; de la Salle, P.; Perillo, M.; Reynolds, J.; Reynolds, R.; Waldron, J. F. (2024). "The last giants: New evidence for giant Late Triassic (Rhaetian) ichthyosaurs from the UK". PLOS ONE. 19 (4). e0300289. doi:10.1371/journal.pone.0300289. PMC 11023487. PMID 38630678.
- ^ Saint-Hilaire, G.É-F. (1831). Recherches sur de grands sauriens trouvés à l’état fossile vers les confins maritimes de la Basse-Normandie, attribués d’abord aux crocodile puis déterminés sous les noms de Teleosaurus et Steneosaurus. Paris: Firmin Didot. pp. 1–138.
- ^ Young, Mark T.; Wilberg, Eric W.; Johnson, Michela M.; Herrera, Yanina; De Andrade, Marco Brandalise; Brignon, Arnaud; Sachs, Sven; Abel, Pascal; Foffa, Davide; Fernández, Marta S.; Vignaud, Patrick; Cowgill, Thomas; Brusatte, Stephen L. (2024). "The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)". Zoological Journal of the Linnean Society. 200 (2): 547–617. doi:10.1093/zoolinnean/zlad165.
- ^ Lydekker, R. (1889). "On the Remains and Affinities of five Genera of Mesozoic Reptiles". Quarterly Journal of the Geological Society. 45 (1–4): 41–59. doi:10.1144/GSL.JGS.1889.045.01-04.04. S2CID 128586645.
- ^ Lee, Hang-Jae (2005). A New Protosuchian (Archosauria: Crocodyliformes) Skull from the Hasandong Formation (Lower Cretaceous) of Hadong, Korea (PDF) (MSc thesis). Chungnam National University.
- ^ Trevor H. Rempert; Brennan P. Martens; Alexander P. M. Vinkeles Melchers (2022). "First Record of a Tylosaurine Mosasaur from the Latest Cretaceous Phosphates of Morocco". Open Journal of Geology. 12 (11): 883–906.
- ^ List of predatory publishers by scholarlyoa.com
- ^ "Spears: 'Universe is Like Space Ship' — and the problem with 'predatory' science journals". Ottawa Citizen. May 16, 2015. Retrieved August 9, 2019.
- ^ Bardet, Nathalie; Fischer, Valentin; Jalil, Nour-Eddine; Khaldoune, Fatima; Yazami, Oussama Khadiri; Pereda-Suberbiola, Xabier; Longrich, Nicholas (February 2025). "Mosasaurids Bare the Teeth: An Extraordinary Ecological Disparity in the Phosphates of Morocco Just Prior to the K/Pg Crisis". Diversity. 17 (2): 114. doi:10.3390/d17020114. ISSN 1424-2818.
- ^ Malkani M.S. (2014). Theropod dinosaurs and mesoeucrocodiles from the Terminal Cretaceous of Pakistan. Abstract Volume of 2nd symposium of International Geoscience Program 608 (IGCP 608) “Cretaceous Ecosystem of Asia and Pacific”. Tokyo, Japan. pp. 169–172.
- ^ Malkani, M. Sadiq (2015). Terrestrial mesoeucrocodiles from the Cretaceous of Pakistan. 12th Symposium on "Mesozoic Terrestrial Ecosystems". pp. 242–246.
- ^ Buchy, Marie-Céline; Frey, Eberhard; Stinnesbeck, Wolfgang; López-Oliva, José Guadalupe (2003). "First occurrence of a gigantic pliosaurid plesiosaur in the late Jurassic (Kimmeridgian) of Mexico" (PDF). Bulletin de la Société géologique de France. 174 (3): 271–278. doi:10.2113/174.3.271. ISSN 1777-5817. S2CID 56469492. Archived from the original (PDF) on March 17, 2024.
- ^ Spindler, Frederik; Mattes, Martin (2021). "Einsame Spitze – Fakten und Spekulation über das Solnhofener Gipfelraubtier" [Lonely at the top – facts and rumours about the Solnhofen apex predator]. Archaeopteryx (in German). 37: 56–68.
- ^ Nessov, L. A. (1990). "Late Jurassic labyrinthodont (Amphibia, Labyrinthodontia) among other relict vertebrate groups in northern Fergana". Paleontologicheskii Zhurnal. 3: 82–90.
- ^ a b Averianov, A. O. (2000). "Sunosuchus sp. (Crocodylomorpha, Goniopholididae) from the Middle Jurassic of Kirghisia" (PDF). Journal of Vertebrate Paleontology. 20 (4): 776–779. doi:10.1671/0272-4634(2000)020[0776:SSCGFT]2.0.CO;2. S2CID 85976137.
- ^ a b Nessov, L. A.; Kaznyshkina, L. F.; Cherepanov, G. O. (1989). "Mesozoic dinosaurians-ceratopses and crocodiles of Central Asia". In Bogdanova, T. N.; Khozatsky, L. I. (eds.). Theoretical and Applied Aspects of Modern Paleontology. Leningrad: Nauka. pp. 144–154.
- ^ Heere, Jelle J.A.; Wallaard, Jonathan J.W.; Mulder, Eric W.A.; Ponstein, J.; Schulp, Anne S. (2023). "The first report of Chelonioidea cf. Ctenochelys from the Late Cretaceous of the Maastrichtian type area". Netherlands Journal of Geosciences. 102. e6. doi:10.1017/njg.2023.3.
- ^ a b Atanassov, M.N. (2002). Two new archosaur reptiles from the Late Triassic of Texas (PhD thesis). Lubbock, Texas: Texas Tech University. pp. 1–352.
- ^ a b Martz, J. W. (2008). Lithostratigraphy, chemostratigraphy, and vertebrate biostratigraphy of the Dockum Group (Upper Triassic), of southern Garza County, West Texas (Unpublished PhD thesis). Texas Tech University. Retrieved 16 November 2021.
- ^ Evans, Mark (2012). A New Genus of Plesiosaur (Reptilia: Sauropterygia) from the Pliensbachian (Early Jurassic) of England, and a Phylogeny of the Plesiosauria (PhD thesis). University of Leicester. pp. 1–397.
- ^ Kear, Benjamin P. (2016). "Cretaceous marine amniotes of Australia: perspectives on a decade of new research" (PDF). Memoirs of Museum Victoria. 74: 17–28. doi:10.24199/j.mmv.2016.74.03.
- ^ Segkar, A. (2022-08-22). "More than just a pretty Penny for the small town of Richmond". ABC Listen. Retrieved 2025-03-06.
- ^ Ritchie, A. (1990). "Return of the Great Sea Monsters". Australian Natural History. 23 (7): 538–545.
- ^ V. Fischer; R. B. J. Benson; P. S. Druckenmiller; H. F. Ketchum; N. Bardet (2018). "The evolutionary history of polycotylid plesiosaurians". Royal Society Open Science. 5 (3): 172177. Bibcode:2018RSOS....572177F. doi:10.1098/rsos.172177. PMC 5882735. PMID 29657811.
- ^ 宇都宮, 聡 (2019-03-31). 鹿児島県長島町獅子島の上部白亜系御所浦層群から産出した 東アジア最古のエラスモサウルス科(爬虫綱,長頚竜目)(Oldest Elasmosauridae (Plesiosauria) in East Asia from the Upper Cretaceous Goshoura Group, Shishijima Island, Southwestern Japan) (Thesis) (in Japanese). Osaka Museum of Natural History.
- ^ "2004 年に鹿児島県長島町獅子島でクビナガリュウの化石が発見 (The finding of plesiosaur fossil from Shishijima, Nagashima-cho, Kagoshima prefecture in 2004)" (PDF). 鹿児島県立博物館 鹿博だより No.101. Kagoshima Prefectural Museum. March 20, 2020. Retrieved August 11, 2023.
- ^ "1億年前の首長竜に現代の鳥に似た習性 ─ 世界初、未消化で吐き出した"ペリット"を発見 ─|東京都市大学". www.tcu.ac.jp. Retrieved 2024-02-14.
- ^ Seeley, H.G. (1869). Index to the fossil remains of Aves, Ornithosauria and Reptilia, from the Secondary system of strata arranged in the Woodwardian Museum of the University of Cambridge. Cambridge, 143 pp.
- ^ Welles, S.P. (1962). "A new species of elasmosaur from the Aptian of Colombia and a review of the Cretaceous plesiosaurs" (PDF). University of California Publications in Geological Sciences. 44 (1): 1–96. ISBN 978-0-598-20148-5. OCLC 5734397. Archived from the original (PDF) on 2022-01-04.
- ^ Malkani, Muhammad Sadiq (2010). "Updated Stratigraphy and Mineral Potential of Sulaiman (Middle Indus) Basin, Pakistan" (PDF). Sindh University Research Journal (Science Series). 42 (2): 39–66.
- ^ Garvey, S.T. (2020). A new high-latitude Tylosaurus (Squamata, Mosasauridae) from Canada with unique dentition (MS). University of Alberta. Archived from the original on 2020-07-23.
- ^ Malkani, Muhammad Sadiq (November 2019). "Recently Discovered Basilosaurid, Baluchithere Rhinoceros, Horses, Sea Cow, Proboscidean, Eucrocodile, Pterosaurs, Plesiosaur, Fishes, Invertebrates and Wood Fossils, Tracks and Trackways of Dinosaurs from Pakistan; Comparison of Recognized Four Titanosaur Taxa of Indo-Pakistan with Madagascar". Open Journal of Geology. 9 (12): 919–955. doi:10.4236/ojg.2019.912098.