Non-avian dinosaurs named from the Chorrillo Formation

The Chorrillo Formation, also named as Chorillo Formation,[1] is a Maastrichtian (Late Cretaceous Epoch, 72.1 - 66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than 50 metres (160 ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation.[2]

Fossils

Dinosaur remains are among the fossils that have been recovered from the formation.

Fossils recovered from the formation are:

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Dinosaurs

Ornithopods

Ornithopods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Euiguanodontia Indet. Indeterminate Proximal end of a femur
Hadrosauridae Indet. Indeterminate Two incomplete caudal centra
Isasicursor[3] I. santacrucensis Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbs An elasmarian ornithopod framless

Sauropods

Sauropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Nullotitan[3] N. glaciaris Estancia Alta Vista A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle bone A colossosaurian titanosaur framless
Titanosauria Indet. Indeterminate Seven isolated teeth

Theropods

Theropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Kookne[3] K. yeutensis An incomplete right coracoid lacking sternal end and proximal end damaged An ornithuran bird
Maip[4] M. macrothorax La Anita Farm Axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal The largest megaraptorid theropod of South America and possibly the largest member of this group

Megaraptoridae Indet. Indeterminate La Anita Farm Isolated teeth
Noasauridae Indet. Indeterminate Isolated right pedal phalanx IV-2?
Unenlagiidae Indet. Indeterminate An isolated dorsal vertebrae, pedal ungual of digit II and phalanx III-2
Yatenavis[5] Y. ieujensis Estancia La Anita. Distal right humerus An enantiornithine bird

Thyreophorans

Thyreophorans of the Chorrillo Formation
Genus Species Locality Material Notes Images
Parankylosauria Indet.[6] Indeterminate Five fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms[7]

Anurans

Amphibians of the Chorrillo Formation
Genus Species Locality Material Notes Images
Anura Indet. Indeterminate Isolated tibiofibula
Calyptocephalella C. gayi Incomplete left maxilla, incomplete right maxilla, incomplete right ilium, proximal right radioulna, left proximal end of tibiofibula and proximal half of urostyle
Calyptocephalellidae Indet. Indeterminate A distal end of right humerus
Pipoidea Indet. Indeterminate Distal end of right humerus

Fishes

Fishes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Lamniformes Indet. Indeterminate Isolated upper lateral tooth
Teleostei Indet. Indeterminate Three crushing tooth crowns
Vidalamiinae Indet. Indeterminate Isolated tooth A amiid fish

Mammals

Mammals of the Chorrillo Formation
Genus Species Locality Material Notes Images
Magallanodon[8] M. baikashkenke Río de Las Chinas Valley and La Anita Farm Tentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right side A ferugliotheriid gondwanathere
Mammalia Indet. Indeterminate Two isolated caudal vertebrae
Patagorhynchus[9] P. pascuali La Anita Farm Lower right molar attached to a fragment of the dentary A ornithorhynchid monotreme
Patagomaia[10] P. chainko La Anita Farm Partial left acetabulum and ischium A large therian

Reptiles

Mosasaurs

Mosasaurs of the Chorrillo Formation
Genus Species Locality Material Notes Images
Mosasauridae Indeterminate

Rhynchocephalian

Rhynchocephalians of the Chorrillo Formation
Genus Species Locality Material Notes Images
Notosphenos[11] N. finisterre A sphenodontid rhynchocephalian

Snakes

Snakes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Rionegrophis R. madtsoioides A partial vertebra. A madtsoiid snake
Serpentes Indeterminate Partial mid-posterior vertebrae

Turtles

Turtles of the Chorrillo Formation
Genus Species Locality Material Notes Images
Chelidae Indet. Indeterminate
Hydromedusinae Indet. Indeterminate Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment
Meiolaniformes Indet. Indeterminate Distal end of a left humerus

Insects

Insects of the Chorrillo Formation
Genus Species Locality Material Notes Images
Coelolepida Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-3:W34/1, MPM-Pal 21835-23:W55/0, MPM-Pal 21835-22:R52/2,MPM-Pal 21835-20:Z47/3. MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3 A Glossatan Lepidopteran
Diamesinae Indet.[12] "Morphotype 3" La Anita farm MPM-Pal 21835-9:E34/4, MPM-Pal 21835-19: V34/4 A Chironomidae Nematoceran. Similar to the living genus Paraheptagyia
Ephemeroptera Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-9:W37/1 A Mayfly
Lepidoptera Indet.[12] Indeterminate La Anita farm MPM-Pal 21835-10:X30/2, MPM-Pal 21835-7:S27/2
Orthocladiinae Indet.[12] "Morphotype 1" La Anita farm MPM-Pal 21835-24: M31/0, MPM-Pal 21835-3: P46/4 A Chironomidae Nematoceran
"Morphotype 2" La Anita farm MPM-Pal 21835-25: V26/3
"Morphotype 5" La Anita farm MPM-Pal 21835-9: Q58/3 It resembles modern Cricotopus
Tanypodinae Indet.[12] "Morphotype 4" La Anita farm MPM-Pal 21835-2:Y40/3 A Chironomidae Nematoceran

Molluscs

Gastropods

Gastropods of the Chorrillo Formation
Genus Species Locality Material Notes Images
Allopeas[13] A. agnolini A achatinid gastropod
Bocourtia[13] B. leonardodavincii A bulimulid gastropod
Lilloiconcha[13] L. novasi A charopid gastropod
Coelostemma[13] C. patagonica A holospirid gastropod
Holospira H. sp.
Physa P. sp.
Pleuroceridae Indet. Indeterminate
Pomacea P. sp.
Potamolithus[13] P. fabiani A tateid gastropod
Stenophysa S. sp.

Plants

Plants of the Chorrillo Formation
Genus Species Locality Material Notes Images
Podocarpoxylon P. dusenii

Algaes

Algaes of the Chorrillo Formation
Genus Species Locality Material Notes Images
Sphaeroplea[14] S. striatocristata

Pollen

Pollen of the Chorrillo Formation
Genus Species Locality Material Notes Images
Acylomurus[14] A. silviae
Baculatisporites B. comaumensis
B. kachaikensis
Ceratosporites C. equalis
Clavatipollenites C. sp.
Clavifera C. triplex
Concavissimisporites C. sp.
Cyatheacidites C. annulatus
Cyathidites C. australis
C. minor
C. rafaeli
Foveosporites F. canalis
Gleicheniidites G. senonicus
Laevigatosporites L. ovatus
Leptolepidites L. sp.
Lusatisporis[14] L. choiols
Microcachryidites M. antarcticus
Peninsulapollis P. gilli
Phyllocladidites P. mawsonii
Podocarpidites cf. P. ellipticus
cf. P. herbstii
P. sp.
Retitriletes R. austroclavatidites
Trichotomosulcites T. subgranulatus
Trilites T. fasolae
cf. T. tuberculiformis
cf. T parvallatus
Tricolpites T. reticulatus
T. sp.
Tuberculatosporites T. parvus
Vitreisporites V. signatus

See also

References

  1. ^ Río Leona at Fossilworks.org
  2. ^ Marenssi et al., 2003, p.406
  3. ^ a b c Fernando Novas; Federico Agnolin; Sebastián Rozadilla; Alexis Aranciaga-Rolando; Federico Brissón-Eli; Matias Motta; Mauricio Cerroni; Martín Ezcurra; Agustin Martinelli; Julia D'Angelo; Gerardo Álvarez-Herrera; Adriel Gentil; Sergio Bogan; Nicolas Chimento; Jordi García-Marsà; Gastón Lo Coco; Sergio Miquel; Fatima Brito; Ezequiel Vera; Valeria Loinaze; Mariela Fernandez; Leonardo Salgado (2019). "Paleontological discoveries in the Chorrillo Formation (upper Campanian-lower Maastrichtian, Upper Cretaceous), Santa Cruz Province, Patagonia, Argentina". Revista del Museo Argentino de Ciencias Naturales. Nueva Series. 21 (2): 217–293. doi:10.22179/revmacn.21.655. hdl:11336/122229.
  4. ^ Aranciaga Rolando AM, Motta MJ, Agnolín FL, Manabe M, Tsuihiji T, Novas FE (2022). "A large Megaraptoridae (Theropoda: Coelurosauria) from Upper Cretaceous (Maastrichtian) of Patagonia, Argentina". Scientific Reports. 12 (1): Article number 6318. Bibcode:2022NatSR..12.6318A. doi:10.1038/s41598-022-09272-z. PMC 9042913. PMID 35474310.
  5. ^ Herrera, Gerardo Álvarez; Agnolín, Federico; Rozadilla, Sebastián; Lo Coco, Gastón E.; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (2022-12-16). "New enantiornithine bird from the uppermost Cretaceous (Maastrichtian) of southern Patagonia, Argentina". Cretaceous Research. 144: 105452. doi:10.1016/j.cretres.2022.105452. ISSN 0195-6671. S2CID 254804249.
  6. ^ Perez Loinaze, Valeria S.; Vera, Ezequiel I.; Raigemborn, M. Sol; Moyano-Paz, Damián; Santamarina, Patricio; Agnolín, Federico; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (2025-01-25). "Palynofloras from the Maastrichtian Chorrillo Formation (Patagonia, Argentina): Paleoenvironmental and paleoclimatic implications". Palaeogeography, Palaeoclimatology, Palaeoecology (in press): 112764. doi:10.1016/j.palaeo.2025.112764. ISSN 0031-0182.
  7. ^ Rozadilla S, Agnolín F, Manabe M, Tsuihiji T, Novas FE (2021). "Ornithischian remains from the Chorrillo Formation (Upper Cretaceous), southern Patagonia, Argentina, and their Implications on ornithischian paleobiogeography in the southern hemisphere". Cretaceous Research. 125: Article 104881. Bibcode:2021CrRes.12504881R. doi:10.1016/j.cretres.2021.104881.
  8. ^ Goin, Francisco J.; et al. (2020). "First Mesozoic Mammal from Chile: the southernmost RECORD of a Late Cretaceous Gondwanatherian" (PDF). Boletín del Museo Nacional de Historia Natural, Chile. 69 (1): 5–31. doi:10.54830/bmnhn.v69.n1.2020.8. S2CID 255652030.
  9. ^ Chimento, N. R.; Agnolín, F. L.; Manabe, M.; Tsuihiji, T.; Rich, T. H.; Vickers-Rich, P.; Novas, F. E. (2023). "First monotreme from the Late Cretaceous of South America". Communications Biology. 6 (1). 146. doi:10.1038/s42003-023-04498-7. PMC 9935847. PMID 36797304.
  10. ^ Chimento, Nicolás R.; Agnolín, Federico L.; García-Marsà, Jordi; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (February 3, 2024). "A large therian mammal from the Late Cretaceous of South America". Scientific Reports. 14 (1): 2854. Bibcode:2024NatSR..14.2854C. doi:10.1038/s41598-024-53156-3. PMC 10838296. PMID 38310138.
  11. ^ Agnolín, Federico; Aranciaga Rolando, Mauro; Manabe, Makoto; Tsuihiji, Takanobu; Novas, Fernando E. (2024-05-02). "Southernmost lepidosaur (Reptilia) assemblage from the Late Cretaceous of Patagonia". Historical Biology: 1–15. doi:10.1080/08912963.2024.2341850. ISSN 0891-2963.
  12. ^ a b c d e f Vera, Ezequiel I.; Monferran, Mateo D.; Massaferro, Julieta; Sabater, Lara M.; Gallego, Oscar F.; Perez Loinaze, Valeria S.; Moyano-Paz, Damián; Agnolín, Federico L.; Manabe, Makoto; Tsuhiji, Takanobu; Novas, Fernando E. (2023-12-11). "A Maastrichtian insect assemblage from Patagonia sheds light on arthropod diversity previous to the K/Pg event". Communications Biology. 6 (1). doi:10.1038/s42003-023-05596-2. ISSN 2399-3642. PMC 10711029.
  13. ^ a b c d e Miquel, S. E. (2024). "New species and records of freshwater and land gastropods from the Cretaceous of Argentina". Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen. 310 (2): 119–131. doi:10.1127/njgpa/2023/1173.
  14. ^ a b c Perez Loinaze, V. S.; Vera, E. I.; Moyano-Paz, D.; Coronel, M. D.; Manabe, M.; Tsuihiji, T.; Novas, F. E. (2023). "Maastrichtian palynological assemblages from the Chorrillo Formation, Patagonia, Argentina". Review of Palaeobotany and Palynology. 314. 104893. Bibcode:2023RPaPa.31404893P. doi:10.1016/j.revpalbo.2023.104893. S2CID 258043990.

Bibliography

Further reading

  • J. E. Powell. 2003. Revision of South American titanosaurid dinosaurs: palaeobiological, palaeobiogeographical and phylogenetic aspects. Records of the Queen Victoria Museum Launceston 111:1-173
Allikas: http://en.wikipedia.org/wiki/Chorrillo_Formation
No tags for this post.