Nemegt Formation

Nemegt Formation
Stratigraphic range: Maastrichtian Possibly also late Campanian
Nemegt, the type locality of the Nemegt Formation
TypeGeological formation
Sub-unitsLower, Middle and Upper units
UnderliesAlluvium
OverliesBaruungoyot Formation
ThicknessOver 235 m (771 ft)
Lithology
PrimaryShale, sandstone
OtherMudstone, conglomerate
Location
Coordinates43°30′N 101°00′E / 43.5°N 101.0°E / 43.5; 101.0
Approximate paleocoordinates40°48′N 90°12′E / 40.8°N 90.2°E / 40.8; 90.2
RegionBayankhongor Aimag, Omnogov, Ovorkhangai
CountryMongolia
ExtentGobi Desert
Nemegt Formation is located in Mongolia
Nemegt Formation
Nemegt Formation (Mongolia)

The Nemegt Formation (also known as Nemegtskaya Svita) is a geological formation in the Gobi Desert of Mongolia, dating to the Late Cretaceous. The formation consists of river channel sediments and contains fossils of fish, turtles, crocodilians, and a diverse fauna of dinosaurs, including birds.

Description

Cretaceous-aged dinosaur fossil localities of Mongolia. Nemegt localities at area A.

The Nemegt Formation is composed of mudstones and sandstones that were deposited by ancient lakes, streams, and flood plains. The Altan Uul locality was described by Michael Novacek as "a canyon carved out of a very rich series of sedimentary rocks" with "steep cliffs and narrow washes". The climate associated with it was wetter than when preceding formations were deposited; there seems to have existed at least some degree of forest cover as indicated by petrified fossil trunks. When examined, the rock facies of the upper section consist of braided meanders, and oxbow lakes. Alteration of sediment grades indicates frequent hydrodynamic condition changes (seasonal rain). Siltstone pellets indicate desiccation at the channel margins in river low-stand periods. Bank erosion features would form through river channel migration. Generally rivers consisted of low flow regimes with sporadic upper flow regimes indicated by exotic pebbles. The lower sections of the formation may represent lacustrine or palustrine deposition.[1] A study of oxygen-18 isotopes from theropod teeth show seasonal patterns similar to present day continental climates. Mean annual temperatures are estimated to be around 7.6 °C (45.7 °F) with seasonal precipitation patterns compared to meteorological data from Shijiazhuang, China; suggesting a monsoonal, cold semi-arid climate.[2] That same study indicated that due to the greenhouse conditions of the Cretaceous, the East Asian monsoon was strong enough to reach into Mongolia. The Nemegt environment was dominated by araucarian conifers,[2] but it also supported a diverse array of other plants, including ginkgos and a wide variety of angiosperms—ranging from plane trees (sycamores) to duckweeds.[3] The groundcover included quillwort like plants as well as sedges (members of the group Cyperaceae).[3] Grass is also known from the formation, and as in the Laramie Formation of North America,[4] reed grasses (primarily Phragmites) were present.[5] The area also supported Bald Cypress.[5] The area probably resembled the modern day Okavango Delta in Botswana.[6]

Stratigraphy

Nemegt and Baruungoyot formations intersection

The most recent stratigraphy divides the Nemegt Formation into three informal members. The lower member is dominated by fluvial deposits, while middle and upper members consist of alluvial plain, paludal, lacustrine, and fluvial deposits. The colour of the sediments is usually light grey to tan in colour in comparison to the typically red colour of the underlying Baruungoyot Formation.[7] It overlies and sometimes interfingers with the Baruungoyot Formation. Interfingering has been noted at the stratotype (Red Walls) and Hermiin Tsav.[7] There has been no absolute dating of the Nemegt Formation. Historically the Nemegt has been considered late Campanian to Maastrichtian, based on comparisons of fossils present, but no exact dating has been performed. U-Pb dating from teeth of Tarbosaurus suggested that the deposition of the middle Nemegt Formation happened before 66.7 ± 2.5 Ma, during the Maastrichtian age, though whether the depositional age of the formation dates back to the late Campanian remains unresolved.[8]

While it has been generally assumed that the Djadochta, Baruungoyot, and Nemegt formations were layered in succession, some geological analysis suggested interfingering rather than superposition of these features. Thus these formations could represent different parts of a single interconnected ecosystem, with the Djadochta reflecting arid dune habitats dominated by small animals, the Baruungoyot representing transitional semi-arid zones, and the Nemegt capturing wetter river plains that supported large dinosaurs.[9]

Paleobiota of the Nemegt Formation

Stratigraphic positions are based on Eberth (2018) who correlated localities to their approximate position within the formation.[7]

Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Flora

Genus Species Location Stratigraphic position Material Notes Image
Araucariaceae[2] Indeterminate Altan Ula II "Petrified wood and trunks" Araucarian conifer, probably the dominant flora of the formation.[2]
Arundo A. sp. Nogon Tsav "Leaf impressions."[5] Cane Grass.
Carpinus C. altanensis Atlan Uul "Fruits."[5] A Hornbeam birch
Cyperaceae indet. Indeterminate Ulaan Bulag "Scattered stem and leaf impressions."[10] A grass-like monocot.
Ginkgo G. altanensis Altan Uul "Leaf impressions."[5] Ginkgophyte.
G. sp. Altan Uul "Leaf impressions."[3] Ginkgophyte.
Lemna L. chushuensis Tsagan Khushu "Fruit."[5] Duckweed.
Lemnaceae indet. Indeterminate Tsagan Khushu, Ulaan Bulag "Fruits."[11] Duckweed.
Monilitheca M. minuta Ulaan Bulag "Two isolated megasporophylls and megaspore casts."[10] A sporophyll related to quillworts.
Nelumbo N. nogontzavensis Nogon Tsav "Leaf and fruit impressions."[5] Water Lotus.
N. sp. Atlan Uul "Leaf impressions."[3] Water Lotus.
Nilssonia N. sp. Atlan Uul "Leaf impressions."[5] Leaves assigned to Cycadophyta or their own order, Nilssoniales.
Nyssoidea N. mongolica Bugiin Tsav Middle or Upper "Fruits." Likely not in the Nyssaceae family. Also in the Bayanshiree formation.[12][13]
Phragmites P. sp. Atlan Uul, Nogon Tsav "Leaf and stem impressions."[5] Reed Grass.
Platanus P. sp. Altan Uul "Leaf impressions."[3] Sycamore.
Potamogeton-like Indeterminate Ulaan Bulag "Shoots, leaves, and fruits."[11] Pondweed.
Silicified Wood Indeterminate Altan Uul II, Nemegt "Petrified wood and trunks."[1] Indeterminate fossil trees.
Taxodium T. sp. Paleocne border locality, likely Tsagan Khushu "Leaf impressions."[5] Cypress.
Trochodendroides T. sp. Altan Uul, and Paleocne border locality, likely Tsagan Khushu "Leaf impressions."[3] A Katsura relative.
Quereuxia Q. sp. Atlan Uul,[3] Nogon Tsav"[5] "Leaf impressions."[3] A Water caltrop look-alike.
Zingiberopsis Z. sp. Nogon Tsav "Leaf impressions."[5] Ginger relative.

Invertebrates

Genus Species Location Stratigraphic position Material Notes Image
Altanicypris A. bispinifera Altan Uul IV, Nemegt Lower "Complete carapace and left valve."[14] An ostracod.
A. multispina Altan Uul IV, Nemegt Lower "Nine carapaces and some detached valves."[14] An ostracod.
A. szczechurae Altan Uul IV, Nemegt Lower "Multiple carapaces and detached valves."[14] An ostracod.
Candona C. altanulaensis Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
cf. C. fabaeformis Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Candoniella C. altanica Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
C. mordvilkoi Altan Uul IV Lower "Three adult carapaces."[15] An ostracod.
Cyclocypris C. transitoria Bügiin Tsav, Nemegt "Seven adult carapaces."[14] An ostracod.
Cypria C. elata Bügiin Tsav, Hermiin Tsav "Several carapaces and valves."[14] An ostracod.
Cypridopsis? C. bugintsavicus Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Ten adult carapaces."[14] An ostracod.
Indeterminate Nemegt Lower "Several complete carapaces."[14] An ostracod.
Cypris? C. ectypa Altan Uul IV, Bügiin Tsav, Nemegt, Ulan Bulag "Complete carapaces and some detached valves."[14] An ostracod.
Cypridea cf. C. punctilataeformis Nemegt "One adult carapace."[15] An ostracod.
C. barsboldi Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
C. cavernosa Altan Uul IV, Bügiin Tsav, Nemegt, Nogon Tsav "Multiple specimens with carapaces and valves."[14] An ostracod.
C. rostrata Bügiin Tsav, Nemegt, Nogon Tsav "Caparaces."[15] An ostracod.
Indeterminate Altan Uul IV, Nemegt Lower "Five juvenile carapaces."[14] An ostracod.
Gobiella G. prima Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Khandia K. stankevitchae Altan Uul IV, Nemegt Lower "Multiple carapaces and valves."[14] An ostracod.
Leiria Indeterminate Nemegt Lower "Several detached valves."[14] An ostracod.
Indeterminate Nemegt Lower "Complete carapace."[14] An ostracod.
Limnocythere sp. Indeterminate Nemegt Lower "Juvenile carapace."[14] An ostracod.
Lycopterocypris? cf. L. profunda Altan Uul IV, Nemegt Lower "Multiple complete carapaces and left valve."[14] An ostracod.
Mongolianella M. palmosa? Altan Uul IV, Bügiin Tsav, Nemegt "Several adult carapaces."[14] An ostracod.
Mongolocypris M. distributa Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Multiple specimens with complete carapaces and valves."[14] An ostracod.
Nemegtia N. biformata Altan Uul IV, Bügiin Tsav, Nemegt, Tsagan Khushu "Multiple complete carapaces and several detached valves."[14] An ostracod.
N. obliquecostae Altan Uul IV, Nemegt Lower "Multiple carapaces and valves."[14] An ostracod.
N. reticulata Altan Uul IV, Bügiin Tsav, Nemegt, Nogon Tsav, Tsagan Khushu "Several carapaces and detached valves."[14] An ostracod.
Paracypridea? P. mongolica Altan Uul IV, Nemegt Lower "Multiple carapaces and detached valves."[14] An ostracod.
Rhinocypris sp. Indeterminate Nemegt Lower "Several carapaces."[14] An ostracod.
Scabriculocypris S. ingenicus? Altan Uul IV, Bügiin Tsav, Naran Bulag, Nemegt, Tsagan Khushu "Multiple carapaces and valves."[14] An ostracod.
S. rasilis? Altan Uul IV, Nemegt Lower "Several carapaces and detached valves."[14] An ostracod.
Timiriasevia cf. T. miaogouensis Altan Uul IV, Nemegt Lower "Seven valves."[14] An ostracod.
cf. T. opinabilis Altan Uul IV Lower "Adult carapace."[15] An ostracod.
T. minuscula Altan Uul IV, Bügiin Tsav, Nemegt "Multiple specimens with carapaces and valves."[14] An ostracod.
T. naranbulakensis Altan Uul IV, Bügiin Tsav, Nemegt "Four carapaces and one right valve."[14] An ostracod.
Ziziphocypris Z. costata Altan Uul IV, Bügiin Tsav, Nemegt "Fourteen carapaces."[14] An ostracod.

Fish

Genus Species Location Stratigraphic position Material Notes Image
Harenaichthys H. lui Altan Uul, Bügiin Tsav, Hermiin Tsav, Ulan Khushu "Partial skull parts, isolated and articulated centra, and articulated caudal fin."[16] An osteoglossomorph. Includes previously reported material.[17][16]
Osteichthyes indet. Indeterminate Bügiin Tsav "Vertebrae and scales found within a specimen of Deinocheirus''."[18] A bony fish.

Amphibians

Genus Species Location Stratigraphic position Material Notes Image
Altanulia A. alifanovi Altan Uul II Middle or Upper "Partial maxilla."[19] A discoglossid. Its classification has been criticized.[20]

Mammals

Genus Species Location Stratigraphic position Material Notes Image
Buginbaatar[21] B. transaltaiensis[21] Bügiin Tsav Middle or Upper "Fragmentary skull."[21] A multituberculate.
"Gurlin Tsav skull" Indeterminate Middle or Partial skull A mysterious metatherian, possibly a sparassodont.
Cimolomyidae indet. Indeterminate Gurilin Tsav Middle to Upper Humerus Shares affinities to taxa from the Hell Creek Formation. [22]

Crocodylomorphs

Genus Species Location Stratigraphic position Material Notes Image
Paralligator[23] P. gradilifrons[23] Nemegt, Nogon Tsav, Ulaan Bulag Lower "Partially complete skulls with sparse body elements."[23] A paralligatorid.

Pterosaurs

Genus Species Location Stratigraphic position Material Notes Image
Unnamed azhdarchid Indeterminate Guriliin Tsav Middle or Upper "Fragmentary cervical vertebrae."[24] A giant azhdarchid.

Turtles

Genus Species Location Stratigraphic position Material Notes Image
Gobiapalone G. breviplastra Alak Shand Khuduk, Altan Uul I, Bügiin Tsav, Bügiin Tsav II, Nemegt, Ulan Khushu "Shell and body elements from multiple specimens."[25] A trionychid.
Gravemys G. barsboldi Bamba Khuduk, Hermiin Tsav, Ingeni Khobur, Tsagan Khushu "Partial to complete shells."[26][27] A lindholmemydid.

Nanhsiungchelyidae indet.

Indeterminate Hermiin Tsav II Lower "Shell fragment."[28] A nanhsiungchelyid.
Indeterminate Khuren Tsav "Damaged shell."[28] A nanhsiungchelyid.
Indeterminate Nemegt Lower "Partial plastron."[28] A nanhsiungchelyid.
Mongolemys M. elegans Bügiin Tsav, Hermiin Tsav, Tsagan Khushu "Skull, shell and body elements from multiple specimens."[29][30] A lindholmemydid.
Mongolochelys M. efremovi Altan Uul II, Altan Uul III, Bügiin Tsav, Guriliin Tsav, Hermiin Tsav, Nemegt, Tsagan Khushu "Skull, shell and body elements from multiple specimens."[31][32] A sichuanchelyid.
Nemegtemys N. conflata Bügiin Tsav, Nemegt "Partial plastron elements."[25] A trionychid.

"Trionyx"

"T". gilbentuensis Gilbentu "Fragmentary shell."[25] A trionychid.
"T". gobiensis Altan Uul III, Bamba Khuduk, Nemegt, Tsagan Khushu "Fragmentary shell and carapace fragments."[25] A trionychid.
Trionychidae indet. Indeterminate Bamba Khuduk, Bügiin Tsav, Guriliin Tsav, Ingeni Khobur "Shell and body remains."[25] A trionychid. Originally identified as Amyda menneri.[25]

Dinosaurs

Ornithischians

Ankylosaurs
Genus Species Location Stratigraphic position Material Notes Image
"Dyoplosaurus"[33] "D". giganteus[33] Nemegt Lower A series of caudal vertebrae, metatarsals, phalanges, osteoderms, and an undescribed partial tail club.[34] An ankylosaurid now regarded as nomen dubium.
Saichania[35] S. chulsanensis[35] Altan Uul IV Lower Partial vertebrae, tail club, and osteoderms.[36] An ankylosaurid also present in the Baruungoyot Formation.
Tarchia[35] T. teresae[36] Altan Uul IV, Hermiin Tsav I Lower A skull, mandibles, vertebrae, tail club, and other undescribed postcranial material.[36] An ankylosaurid.
T. tumanovae[37] Hermiin Tsav Lower[7] "Partial skeleton with complete skull."[37] An ankylosaurid.
Tetrapodosaurus-like indet. Indeterminate Bügiin Tsav, Shar Tsav "Footprint casts."[38] Ankylosaurid tracks.
Ankylosauridae indet. Indeterminate Altan Uul II-III-IV, Bügiin Tsav, Hermiin Tsav I, Khuree Tsav, Nemegt A partial dentary, caudal vertebrae, dorsal vertebrae, metatarsals, phalanges, pedal phalanx, humeri, partial pelvis, tail clubs, cervical half-rings, osteoderms, skin impressions, and undescribed juveniles and partial skeletons.[34] Cranial and postcranial material representing numerous individuals of different ages.
Hadrosaurs
Genus Species Location Stratigraphic position Material Notes Image
Amblydactylus-like indet. Indeterminate Bügiin Tsav, Bügiin Tsav II, Guriliin Tsav, Yagaan Khovil Middle & Upper[7] "Footprint casts."[38] Three-toed hadrosaurs tracks. Attributed to Saurolophus. Some tracks are huge,those from Guriliin Tsav and Bügiin Tsav up to 1m in length.
Barsboldia[39] B. sicinskii[39] Nemegt Lower[7] "Partial vertebrae, pelvis, and ribs."[39][40] A hadrosaurid.
Hadropodus Indeterminate Nemegt "Numerous footprints."[41] Hadrosaurid tracks. Attributed to Saurolophus.
Saurolophus[42] S. angustirostris[43] Presence in most localities[44] "Multiple specimens, including articulated skulls, postcranial skeletons and mass death assemblage."[45][46][47][48] A saurolophine hadrosaurid which was the most common hadrosaur in the area.[49]
Unnamed ichnotaxon Indeterminate Nemegt "Numerous footprint casts with trackways."[50] Hand and feet hadrosaur tracks. Attributed to Saurolophus.
Pachycephalosaurs
Genus Species Location Stratigraphic position Material Notes Image
Homalocephale[51] H. calathocercos[51] Nemegt Lower[7] "Partial skull and skeleton including hindlimbs."[51] A pachycephalosaurid.
Prenocephale[51] P. prenes[51] Guriliin Tsav, Nemegt "Complete skull and partial postcranium of several specimens."[51][53] A pachycephalosaurid.
P. sp.[53] Tsagan Khushu, Nemegt Lower[7] "complete left squamosal"; large skull
Indeterminate Bugiin Tsav Middle[52] "incomplete frontoparietal dome and fragments of associated braincase bones"[53]

Sauropods

Genus Species Location Stratigraphic position Material Notes Image
Brontopodus Indeterminate Nemegt Lower "Large footprint and several four-toed tracks."[54][41] Sauropod tracks. Tracks were produced by sauropods larger than any in Nemegt known from skeletal remains.[54] Foot measurements vary from 75-90cm in length.[54] Hip height of track maker estimated as 4m.[55]
Brontopodus-like indet. Indeterminate Bügiin Tsav, Yagaan Khovil Middle & Upper [7] "Footprint casts."[38] Sauropod tracks. Footprints from Bügiin Tsav are enormous, up to 85 cm in length.[38]
Nemegtosaurus[56] N. mongoliensis[56] Nemegt Lower "Skulls and body remains from several specimens."[57][58][59] A titanosaur.
Opisthocoelicaudia[60] O. skarzynskii[60] Altan Uul IV Lower "Nearly complete skeleton without the skull and cervical series."[59] A titanosaur.
Undescribed sauropods Indeterminate Altan Uul I-II-II-IV, Bügiin Tsav, Guriliin Tsav, Hermiin Tsav, Khamaryn Khural, Khuree Tsav, Nemegt, Nogon Tsav, Tsagan Khushu, Ulan Khushu "Several fragmentary remains.[59] Sauropod remains that may represent already named taxa,such as Nemegtosaurus.[52]
Unnamed ichnotaxon Indeterminate Nemegt "Footprint casts with skin impressions."[50] Four-toed sauropod tracks. Attributed to Opisthocoelicaudia.

Theropods

Alvarezsaurs
Genus Species Location Stratigraphic position Material Notes Image
Manipulonyx[61] M. reshetovi[61] Hermiin Tsav Lower Partial skeleton including several vertebrae, much of the forelimb, and parts of the hindlimb, pectoral girdle, and pelvic girdle[61] A parvicursorine alvarezsaurid
Mononykus[62] M. olecranus[62] Bügiin Tsav Middle[52] "Fragmentary skull, vertebrae, limbs and fragmented pelvis."[63] An alvarezsaurid
Indeterminate Altan Uul III Lower "Caudal vertebrae and left hindlimb."[64]
Nemegtonykus[64] N. citus[64] Altan Uul III Lower "Partially complete skeleton from two specimens lacking the skull."[64] An alvarezsaurid
Birds
Genus Species Location Stratigraphic position Material Notes Image
Brodavis[65] B. mongoliensis[65] Bügiin Tsav "Nearly complete left tarsometatarsus."[65] A hesperornithine.
Gurilynia[66] G. nessovi[66] Guriliin Tsav Middle[52] "Left coracoid and partial humerii."[66] An enantiornithine.
Judinornis[67] J. nogontsavensis[67] Nogon Tsav "Dorsal vertebra."[67] A hesperornithine.
Laevisoolithus[68] L. sochavai "Whole egg with partly broken pole"[68] Laid by a bird or small theropod.[68]
Subtiliolithus[68] S. microtuberculatus[68] "Eggshell fragments"[68]
Teviornis[69] T. gobiensis[69] Guriliin Tsav Middle[52] "Partial forelimbs."[69] A presbyornithid anseriform.
Unnamed ornithurans Indeterminate Tsagan Khushu "A humerus and two tibiotarsi."[70] Ornithuran remains.
Dromaeosaurs
Genus Species Location Stratigraphic position Material Notes Image
Adasaurus[71] A. mongoliensis[71] Bügiin Tsav Middle[52] "Partial skull and fragmentary postcrania from two specimens."[72] A dromaeosaurid.

Dromaeosauridae indet.

Indeterminate Bügiin Tsav Middle[52] "Cervical vertebra."[73] A dromaeosaurid.
Indeterminate Hermiin Tsav Lower "Dorsal vertebra."[74]
Indeterminate Khaichin I "Three teeth."[75]
Indeterminate Nemegt Lower "Dorsal vertebra and other elements."[74][76]
Indeterminate Indeterminate Bügiin Tsav "Footprints with poor claw marks."[77] Two-toed theropod tracks. Attributed to either Adasaurus or Zanabazar.
Velociraptor?[78] V. sp
Ornithomimosaurs
Genus Species Location Stratigraphic position Material Notes Image
Anserimimus[79] A. planinychus[79] Bügiin Tsav Middle or Upper[7] "Partial skeleton lacking the skull."[79][80] An ornithomimid.
Deinocheirus[81] D. mirificus[81] Altan Uul III-IV, Bügiin Tsav "Complete skull with virtually complete postcranial remains from three specimens."[18] A giant deinocheirid.
Gallimimus[82] G. bullatus[82] Presence in most localities[83] "Multiple specimens with nearly complete skull and postcranial elements."[83][84] An ornithomimid.
Indeterminate Indeterminate Bügiin Tsav "Nine trackways including an articulated Gallimimus foot."[77] Three-toed theropod tracks. Attributed to ornithomimids.
Unnamed ornithomimid[85] Indeterminate Tsagan Khushu "Partial vertebral column with fore and hindlimbs."[85][83] An ornithomimid.
Oviraptorosaurs
Genus Species Location Stratigraphic position Material Notes Image
Avimimus[86] A. portentosus[86] Shar Tsav

Bügiin Tsav[87]

Middle & Upper [7] "Skull roof with fragmentary body elements."[88] An avimimid.
A. nemegtensis[88] Nemegt Lower[7] "Fragmented skull with partial skeleton."[88] An avimimid.
Conchoraptor? C. gracilis? Guriliin Tsav Middle[52] "Skull and a flock of articulated individuals."[89] An oviraptorid. Also present in the Baruungoyot Formation. Possibly representatives of Conchoraptor.
Elmisaurus[90] E. rarus[90] Altan Uul II, Nemegt
  • Lower
  • Middle or Upper
 "Fragmentary skull and partially skeletons from several specimens."[88][91] A caenagnathid.
Elongatoolithidae indet. Indeterminate Bügiin Tsav "Three eggs containing embryonic remains."[92] Oviraptorid eggs.
Gobiraptor[93] G. minutus[93] Altan Uul III Lower "Partial crania with complete lower jaws and fragmented postcrania."[93] An oviraptorid.
Indeterminate Indeterminate Hermiin Tsav, Shar Tsav, Yagaan Khovil "Footprint casts."[38] Three-toed theropod tracks. Attributed to Avimimus.
Nemegtomaia[94] N. barsboldi[94] Nemegt Lower "Multiple specimens including a nesting specimen and eggs."[95] An oviraptorid also present in the Baruungoyot Formation.
Nomingia[96] N. gobiensis[96] Bügiin Tsav, Nemegt "Vertebral series with partial hindlimbs and a referred pygostyle."[88] A caenagnathid. May be synonymous with Elmisaurus.[91]
Oksoko[97] O. avarsan Bügiin Tsav, Guriliin Tsav Middle[52] "Multiple associated skeletons."[97] An oviraptorid.
Rinchenia[98] R. mongoliensis[98] Altan Uul II Middle or Upper "Skull with nearly complete skeleton."[88] An oviraptorid.
Therizinosaurs
Genus Species Location Stratigraphic position Material Notes Image
Therizinosaurus[99] T. cheloniformis[99] Altan Uul II, Hermiin Tsav, Nemegt Lower & Middle[7] "Fore and hindlimb elements from multiple specimens."[100][101] A giant therizinosaurid.
Undescribed therizinosaurids Indeterminate Altan Uul IV, Bügiin Tsav, Tsagan Khushu [102] Lower [7]

Middle & Upper [7]

"Partial remains."[103] Therizinosaurid remains.[102] Possibly Therizinosaurus.
Troodontids
Genus Species Location Stratigraphic position Material Notes Image
Borogovia[104] B. gracilicrus[104] Altan Uul IV Lower "Partial hindlimbs."[104] A troodontid.
Indeterminate Indeterminate Bügiin Tsav "Footprints with poor claw marks."[77] Two-toed theropod tracks. Attributed to either Adasaurus or Zanabazar.
Tochisaurus[105] T. nemegtensis[105] Nemegt Lower "Left metatarsus."[105] A troodontid.
Zanabazar[106] Z. junior[107] Bügiin Tsav Middle[52] "Skull with fragmentary postcranial skeleton."[106] A troodontid originally identified as "Saurornithoides junior".
Tyrannosaurs
Genus Species Location Stratigraphic position Material Notes Image
Alioramus[108] A. remotus[108] Nogon Tsav "Partially preserved skull with very sparse postcranial remains."[108] An alioramin.
A. altai[109] Tsagan Khushu "Nearly complete skull with partial postcrania."[109] An alioramin.
Bagaraatan[110] B. ostromi[110] Nemegt Lower[7] "Fragmentary lower jaw, hindlimb and caudal vertebrae."[110] A tyrannosauroid.
Indeterminate Indeterminate Bügiin Tsav, Bügiin Tsav II "Footprint casts."[38] Three-toed theropod tracks. Attributed to Tarbosaurus.

Raptorex[111]

R. kriegsteni[111]

"Nearly complete skeleton including the skull."[111] A controversial genus of tyrannosaurid that could be a juvenile Tarbosaurus.[112]
Tarbosaurus[113] T. bataar[114] Presence in most localities[2] "Nearly complete skeletons from multiple specimens."[113][114][2] A large tyrannosaurid which was the most common large carnivore in the area.[115]
Unnamed ichnotaxon Indeterminate Nemegt "Footprint casts with skin impressions."[50] Three-toed theropod tracks. Attributed to Tarbosaurus.
Tyrannosauripodidae indet. Indeterminate Nemegt "Five tracks."[41] Three-toed tyrannosaurid tracks. Attributed to either Alioramus or Tarbosaurus.

See also

References

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allikas: https://en.wikipedia.org/wiki/Nemegt_Svita